law of tolerance was given by\tshelford\thaeckel\tJ.grinnel\tgause

The correct option is 1.
Shelford's Law of Tolerance is a principle developed by American zoologist Victor Ernest Shelford in 1911. It states that an organism's success is based on a complex set of conditions and that each individual or population has a certain minimum, maximum, and optimum environmental factor or combination of factors that determine success.